From what Sal said yesterday, the center now has much more defensive responsibilities, covering the low slot when our D chase into the corners. Previously, it would be any forward, often a wing.
Because of this, the center no longer flies the zone as soon as it looks like we get possession, so that results in more support for the D, and shorter breakout passes.
The off wing, however, has the option to go, so they're more playing Jack at wing than C, allowing him to be the deep threat on his line.